1. Department of Plant Biology and Ecology, University of the Basque Country (UPV/EHU), Apdo. 644, E-48080 Bilbao, Spain
2. Departamento de Bioqu�mica Vegetal y Biolog�a Molecular, Facultad de Qu�mica, Universidad de Sevilla, Calle Profesor Garc�a Gonz�lez, 1, Sevilla, Spain
3. Ikerbasque, Basque Foundation for Science, Bilbao, Spain