1. School of Environmental and Life Sciences, University of Newcastle, Callaghan, NSW 2308, Australia
2. Priority Research Centre for Reproductive Science, University of Newcastle, Callaghan, NSW 2308, Australia
3. School of Biological Sciences, University of Auckland, Auckland 1010, New Zealand
4. School of Biomedical Sciences and Pharmacy, University of Newcastle, Callaghan, NSW 2308, Australia