1. Biology Department, Vancouver Island University, Nanaimo, BC, Canada
2. School of Environmental and Rural Studies, University of New England, Armidale, NSW, Australia
3. Forest Genetics Laboratory, Forestry and Forest Products Research Institute, Matsunosato, Ibaraki, Japan
4. Ecosystem Science and Management Program, University of Northern British Columbia, Prince George, BC, Canada
5. Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada