1. Department of Biomedical Sciences, Seoul National University Graduate School , Seoul , Korea (complete list of author affiliations is available in Appendix S1 ; see Supporting Information )
2. Department of Pathology, Seoul National University College of Medicine , Seoul , Korea
3. Department of Dermatology, CHA Bundang Medical Center, CHA University , Seongnam , Korea
4. Department of Dermatology, SMG-SNU Boramae Medical Center , Seoul , Korea
5. Department of Dermatology, Seoul National University College of Medicine , Seoul , Korea
6. Institute of Human-Environment Interface Biology, Medical Research Center, Seoul National University , Seoul , Korea
7. Department of Pathology, CHA Bundang Medical Center, CHA University , Seongnam , Korea
8. Laboratory for Investigative Dermatology, The Rockefeller University , New York, NY , USA
9. Dermatology Section, Veterans Affairs Northern California Health Care System , Mather, CA , USA
10. Department of Dermatology, University of California Davis , Sacramento, CA , USA
11. Department of Microbiology and Immunology, Seoul National University College of Medicine , Seoul , Korea
12. Cancer Research Institute, College of Medicine, Seoul National University , Seoul , Korea
13. Department of Dermatology, Seoul National University Hospital , Seoul , Korea
14. Genomic Medicine Institute, Medical Research Center, Seoul National University , Seoul , Korea