1. School of Anatomical Sciences, Faculty of Health Sciences, University of the Witwatersrand, Parktown, Johannesburg, Republic of South Africa
2. Department of Zoology, KSU Mammals Research Chair, College of Science, King Saud University, Riyadh, Saudi Arabia
3. Centre for Zoo and Wild Animal Health, Copenhagen Zoo, Frederiksberg, Denmark
4. South African Research Chair of Mammal Behavioural Ecology and Physiology, University of Pretoria, Pretoria, South Africa