1. Laboratoire Évolution and Diversification Biologique, Université Paul Sabatier Toulouse III, UMR 5174, CNRS/IRD , 31077 Toulouse , France
2. Department of Biology, University of North Carolina at Chapel Hill , Chapel Hill, NC , United States
3. Department of Evolutionary Biology and Environmental Studies, University of Zurich , Zurich , Switzerland
4. Swiss Data Science Center, ETH , Zurich , Switzerland