1. Department of Anatomy, Physiology and Human Biology, School of Human Sciences, The University of Western Australia , Perth, WA , Australia
2. International Centre of Biodiversity and Primate Conservation, Dali University , Dali , China
3. Centre for Evolutionary Biology, School of Biological Sciences, The University of Western Australia , Perth, WA , Australia
4. Centre of Excellence in Biodiversity and Natural Resource Management, University of Rwanda , Huye , Rwanda
5. Department of Evolutionary Biology and Environmental Studies, University of Zurich , Zurich , Switzerland