1. Department of Biology, Ghent University , Ghent , Belgium
2. Department of Plant Biotechnology and Bioinformatics, Ghent University and VIB-UGent Center for Plant Systems Biology , Ghent , Belgium
3. College of Horticulture, Academy for Advanced Interdisciplinary Studies, Nanjing Agricultural University , Nanjing , China
4. Department of Biochemistry, Genetics and Microbiology, University of Pretoria , Pretoria , South Africa