1. Department of Evolution, Ecology and Organismal Biology, The Ohio State University, 1315 Kinnear Road, Columbus, OH 43212, USA
2. Chicago Botanic Garden, 1000 Lake Cook Road, Glencoe, IL 60022 USA
3. University of Colorado Museum of Natural History, Bruce Curtis Building, UCB 265, University of Colorado, Boulder, CO 80309 USA
4. Department of Ecology and Evolutionary Biology, University of Colorado, Ramaley N122, Campus Box 334, Boulder, CO 80309-0334, USA
5. Department of Zoology, Field Museum of Natural History, 1400 South Lake Shore Drive, Chicago, IL 60605-2496, USA
6. Rancho Santa Ana Botanic Garden, 1500 N College Avenue, Claremont, CA 91711, USA