1. Department of Genetics, Evolution and Environment, University College London, Gower Street, London WC1E 6BT, UK
2. Department of Mathematics, Beijing Jiaotong University, No. 3 Shangyuancun, Haidian District, Beijing, 100044, P.R. China
3. Department of Ecology and Evolutionary Biology, University of Connecticut, 75 N. Eagleville Road, Unit 3043, Storrs, CT 06269-3043, USA
4. Department of Biology & Burke Museum of Natural History and Culture, University of Washington, Seattle, WA 98195-1800, USA