1. Department of Biology & Burke Museum of Natural History and Culture, University of Washington, Seattle, USA
2. National Center for Mathematics and Interdisciplinary Sciences, Academy of Mathematics and Systems Science, Chinese Academy of Sciences, China
3. Key Laboratory of Random Complex Structures and Data Science, Academy of Mathematics and Systems Science, Chinese Academy of Sciences, China
4. Department of Evolution and Ecology, University of California Davis, One Shields Avenue, Davis, USA
5. Department of Genetics, University College London, London, UK
6. Radcliffe Institute for Advanced Studies, Harvard University, Cambridge, USA