1. Division of Life Sciences, Korea Polar Research Institute, 26 Songdomirae-ro, Yeonsu-gu, Incheon 21990, Republic of Korea
2. Unité Bioinformatique Evolutive, Institut Pasteur, Paris, France
3. Institut de Systématique, Evolution, Biodiversité (ISYEB - UMR 7205, CNRS, Muséum National d’Histoire Naturelle, SU, EPHE, UA), Paris, France
4. Departments of Biological Sciences and Statistics, North Carolina State University, Raleigh, NC 27695-7566, USA