1. Plant Ecology and Evolution, Department of Ecology and Genetics, Uppsala University , Norbyvägen 18D, 75236 Uppsala , Sweden
2. Science for Life Laboratory (SciLifeLab), Uppsala University , 75237 Uppsala , Sweden
3. Organismal and Evolutionary Biology Research Program, Faculty of Biological and Environmental Sciences, and Viikki Plant Science Centre, University of Helsinki , P.O. Box 65 (Viikinkaari 1), 00014 Helsinki , Finland
4. School of Biological Sciences, Nanyang Technological University , 60 Nanyang Drive, Singapore 637551 , Singapore