1. Department of Biological Sciences, The George Washington University, 2023 G St. NW, Washington, D.C. 20052, USA; 2College of Fisheries and Life Science, Shanghai Ocean University, Shanghai 201306, China; and 3National Systematics Laboratory NMFS/NOAA, Post Office Box 37012, Smithsonian Institution NHB, WC 60, MRC-153, Washington, D.C. 20013-7012, USA