1. Department of Microbiology and Immunology, Rega Institute, KU Leuven, Leuven, Belgium;
2. Institute of Evolutionary Biology, University of Edinburgh, Edinburgh, United Kingdom;
3. Fogarty International Center, National Institutes of Health, Bethesda, MD, USA;
4. Departments of Biomathematics and Human Genetics, David Geffen School of Medicine at UCLA, University of California, Los Angeles, CA, 90095, USA;
5. Department of Biostatistics, UCLA Fielding School of Public Health, University of California, Los Angeles, CA, 90095, USA