1. Department of Palaeontology and Museum, University of Zurich, Karl-Schmid-Strasse 4, 8006 Zurich, Switzerland
2. Zoological Institute, University of Basel, Vesalgasse 1, 4051 Basel, Switzerland
3. Centre for Ecological and Evolutionary Synthesis (CEES), Department of Biosciences, University of Oslo, 0316 Oslo, Norway