1. Centre for GeoGenetics, Natural History Museum of Denmark
2. Department of Biology, University of Copenhagen, Universitetsparken 15, 2100 Copenhagen Ø, Denmark
3. Department of Geology, University of Otago, PO Box 56, Dunedin 9054, New Zealand
4. Centre for Macroevolution and Macroecology, Research School of Biology, Australian National University, Canberra ACT 0200, Australia
5. Department of Ecology and Evolutionary Biology
6. Fuller Evolutionary Biology Program, Cornell Lab of Ornithology, Cornell University, Ithaca, NY 14850, USA
7. Department of Biology
8. Department of Statistics, University of California, Berkeley, CA 94720-314, USA
9. Center for Macroecology, evolution and Climate, Department of Biology, University of Copenhagen, Universitetsparken 15, 2100 Copenhagen Ø, Denmark