1. Landscape Ecology, Institute of Terrestrial Ecosystems Department of Environmental Systems Sciences, , ETH Zürich, Zurich 8092, Switzerland
2. Swiss Federal Institute for Forest , Snow and Landscape Research WSL, Birmensdorf 8903, Switzerland
3. German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig , Leipzig 04103, Germany
4. Yale University Department of Ecology and Evolutionary Biology, , New Haven, CT 06520, USA
5. Yale University Center for Biodiversity and Global Change, , New Haven, CT 06520, USA