1. School of Life Science, Luoyang Normal University , 6 Jiqing Road, Luoyang 471934 , P. R. China
2. Institute of Ecology and Evolution, The University of Edinburgh , Edinburgh, EH9 3FL , UK
3. Royal Botanic Garden Edinburgh , 20A Inverleith Row, Edinburgh, EH3 5LR , UK
4. Key Laboratory of Adaptation and Evolution of Plateau Biota, Northwest Institute of Plateau Biology, Chinese Academy of Sciences , 810001 Xining , P. R. China