1. Botanical Gardens, Graduate School of Science, The University of Tokyo , 3-7-1 Hakusan, Bunkyo-ku, Tokyo , Japan
2. Faculty of Applied Biological Sciences, Gifu University , Yanagido 1-1, Gifu , Japan
3. Department of Ecology and Evolution, Faculty of Biology and Medicine, University of Lausanne , Lausanne , Switzerland
4. Department of Life Science, Institute of Ecology and Evolutionary Biology, National Taiwan University , Taipei 10617 , Taiwan
5. Chicago Botanic Garden , 1000 Lake Cook Road, Glencoe, IL 60022 , USA