1. Department of Ecology , Evolution, and Behavior, , St. Paul, MN 55108 , United States
2. BioTechnology Institute, University of Minnesota , Evolution, and Behavior, , St. Paul, MN 55108 , United States
3. Division of Community Health & Epidemiology, University of Minnesota School of Public Health , Minneapolis, MN 55454 , United States
4. School of Mathematics, University of Minnesota , Minneapolis, MN 55455 , United States