1. Sustainable Soils and Crops, Rothamsted Research , Harpenden, Hertfordshire AL5 2JQ, United Kingdom
2. Cranfield University School of Water, Energy and Environment, , Cranfield, Bedfordshire MK43 0AL, United Kingdom
3. Swedish University of Agricultural Sciences Department of Ecology, , Uppsala SE-750 07, Sweden
4. INRAE, UR1264 MycSA Present address: , CS2032, 33882 Villenave d’Ornon, France