1. Department of Biology , , Waterloo, ON N2L 3G1 , Canada
2. University of Waterloo , , Waterloo, ON N2L 3G1 , Canada
3. Super-cutting-edge Grand and Advanced Research (SUGAR) Program , Institute for Extra-cutting-edge Science and Technology Avant-garde Research, , Yokosuka, Kanagawa , Japan
4. Japan Agency for Marine-Earth Science and Technology , Institute for Extra-cutting-edge Science and Technology Avant-garde Research, , Yokosuka, Kanagawa , Japan
5. Department of Biological Sciences , , Edmonton, AB T6G 2E9 , Canada
6. University of Alberta , , Edmonton, AB T6G 2E9 , Canada
7. Department of Earth & Environmental Sciences , , Waterloo, ON N2L 3G1 , Canada