1. Department of Oceanography, Texas A&M University, College Station , TX 77843 , United States
2. Department of Marine Chemistry and Geochemistry, Woods Hole Oceanographic Institution , Woods Hole, MA 02543 , United States
3. Biology Department, Carleton College , Northfield, MN 55057 , United States
4. Biology Department, Woods Hole Oceanographic Institution , Woods Hole, MA 02543 , United States
5. Department of Geology & Geophysics, Woods Hole Oceanographic Institution , Woods Hole, MA 02543 , United States