1. Shandong Key Laboratory of Water Pollution Control and Resource Reuse, School of Environmental Science and Engineering, Shandong University , Qingdao 266237 , China
2. Shandong Key Laboratory of Water Pollution Control and Resource Reuse , School of Environmental Science and Engineering, , Qingdao 266237 , China
3. Shandong University , School of Environmental Science and Engineering, , Qingdao 266237 , China
4. State Key Laboratory of Microbial Technology, Microbial Technology Institute, Shandong University , Qingdao, Shandong 266237 , China
5. Department of Biochemistry and Molecular Biology, Pennsylvania State University , University Park, PA 16801 , United States