1. Vanderbilt Genetics Institute and Division of Genetic Medicine , Department of Medicine, Vanderbilt University Medical Center, Nashville, TN 37232 , USA
2. Department of Epidemiology , University of North Carolina, Chapel Hill, NC 27599 , USA
3. The University of Texas Health Science Center at Houston (UTHealth) School of Public Health , Brownsville, TX 78520 , USA
4. Clare Hall , University of Cambridge, Cambridgeshire, UK