1. Section of Ophthalmology , School of Life Course Sciences, , London SE1 7EH, UK
2. King’s College London , School of Life Course Sciences, , London SE1 7EH, UK
3. Department of Twin Research and Genetic Epidemiology , School of Life Course Sciences, , London SE1 7EH, UK
4. NIHR Biomedical Research Centre at Moorfields Eye Hospital NHS Foundation Trust and the UCL Institute of Ophthalmology , London WC1E 6BT, UK
5. Johns Hopkins Bloomberg School of Public Health , Baltimore, MD 21205, USA
6. Department of Ophthalmology , , London SE1 7EH, UK
7. St Thomas’ Hospital, Guy’s and St Thomas’ NHS Foundation Trust , , London SE1 7EH, UK
8. Institute of Ophthalmology , , London WC1E 6BT, UK
9. University College London , , London WC1E 6BT, UK
10. UCL Great Ormond Street Hospital Institute of Child Health , London WC1N 1EH, UK
11. Ulverscroft Vision Research Group , , London WC1N 1EH, UK
12. University College London , , London WC1N 1EH, UK