1. Department of Medicine, University of Cambridge , Level 5, PO Box 157, Cambridge CB2 0QQ , UK
2. Wellcome Trust-Medical Research Council Cambridge Stem Cell Institute, University of Cambridge , Cambridge CB2 0AW , UK
3. Department of Clinical Neuro sciences, University of Cambridge , Cambridge CB2 2PY , UK
4. University of Cambridge, Level 5, PO Box 157 Department of Medicine, , Cambridge CB2 0QQ , UK