1. Faculty of Medicine, School of Biomedical Sciences, The University of Queensland, St Lucia 4072, Australia
2. Institute for Molecular Bioscience, The University of Queensland, St Lucia 4072, Australia
3. Centre for Microscopy and Microanalysis, The University of Queensland, St Lucia 4072, Australia
4. State Key Laboratory of Molecular Developmental Biology, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, China