1. Department of Evolutionary and Environmental Biology, Faculty of Natural Sciences, University of Haifa , 199 Abba Khoushi Ave. Mt. Carmel, Haifa 3498838 , Israel
2. Central Fish Health Laboratory, Fishery and Aquaculture Department, Ministry of Agriculture and Rural Development , 1 Havazelet St. Nir David 1080300 , Israel
3. Department of Biology and Environment, Faculty of Natural Sciences, University of Haifa , Oranim, Derech Kiryat Amal, Tivon 3600600 , Israel