1. Department of Biology, Drexel University , 3245 Chestnut St., Philadelphia, PA 19104, United States
2. State Key Laboratory of Earth Surface Processes and Resource Ecology and Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, College of Life Sciences, Beijing Normal University , Beijing 100875 , China
3. Department of Ecology and Evolutionary Biology, Cornell University , Dale R.Corson Hall, Ithaca, NY 14850, United States
4. Institute of Environmental Sciences, Jagiellonian University , Gołębia 24, 31-007 Kraków, Poland
5. Department of Biology , Calvin College, 1726 Knollcrest Circle SE, Grand Rapids, MI 49546-4402, United States