1. Estación Experimental del Zaidín, CSIC, Departamento de Microbiología del Suelo y Sistemas Simbióticos, Prof. Albareda 1, 18008, Granada, Spain
2. Netherlands Institute of Ecology (NIOO-KNAW), Department of Microbial Ecology, Droevendaalsesteeg 10, 6708 PB, Wageningen, The Netherlands
3. Leiden University, Department of Biology, Leiden, 2311 EZ, The Netherlands
4. Maranhão State University (UEMA), department of Agroecology, São Luís, Brazil