1. International Centre for Terrestrial Antarctic Research, University of Waikato - Te Whare Wānanga o Waikato, Hamilton 3240, New Zealand
2. Centre for Microbial Ecology and Genomics, University of Pretoria, Hatfield 0028, Pretoria, South Africa
3. Institute for Marine and Antarctic Studies, College of Science and Engineering, University of Tasmania, Battery Point, Tasmania, Australia
4. Department of Biochemistry and Microbiology, School of Environmental and Biological Sciences, Rutgers University, New Brunswick, NJ 08901, USA