1. Guangzhou Key Laboratory of Subtropical Biodiversity and Biomonitoring, South China Normal University, Guangzhou 510631, China
2. Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, UK
3. Biodiversity department and Centre for Water and Environmental Research, University of Duisburg-Essen, 45141 Essen, Germany
4. Department of Zoology, South Parks Road, Oxford OX1 3PS
5. Centre for Environment, Fisheries and Aquaculture Science (Cefas), Barrack Road, the Nothe, Weymouth, Dorset DT4 8UB, UK