1. Marine Ecosystem and Biological Research Center, Korea Institute of Ocean Science and Technology, Haeyang-ro, Yeongdo-gu, Busan 49111, Republic of Korea
2. Department of Marine Biology, Korea University of Science and Technology, Gajeong-ro, Yuseong-gu, Daejeon 34113, Republic of Korea
3. Operational Oceanography Research Center, Korea Institute of Ocean Science and Technology, Haeyang-ro, Yeongdo-gu, Busan 49111, Republic of Korea