1. Key Laboratory of Alpine Ecology, Institute of Tibetan Plateau Research, Chinese Academy of Sciences (CAS), Beijing 100101, China
2. College of Resources and Environment, University of Chinese Academy of Sciences, Beijing 100039, China
3. College of Biological and Pharmaceutical Sciences, China Three Gorges University, Yichang 443002, China
4. CAS Center for Excellence in Tibetan Plateau Earth Sciences, Chinese Academy of Sciences, Beijing 100101, China
5. Key Lab Drinking Water Science & Technology, Research Center for Eco-Environmental Sciences, Chinese Academy of Sciences, Beijing 100086, China
6. Key Laboratory of Tibetan Environmental Changes and Land Surface Processes, Institute of Tibetan Plateau Research, Chinese Academy of Sciences, Beijing 100101, China
7. Centre for Microbial Ecology and Genomics, Department of Biochemistry, Genetics and Microbiology, University of Pretoria, Pretoria 0028, South Africa