1. School of Life Science and Technology, Tokyo Institute of Technology, 2-12-1-W3-48 Ookayama, Meguro-ku, Tokyo 152–8550, Japan
2. Center for Information Biology, National Institute of Genetics, 1111 Yata, Mishima, Shizuoka 411–8540, Japan
3. Japan Collection of Microorganisms, RIKEN BioResource Research Center, 3-1-1 Koyada, Tsukuba, Ibaraki 305-0074, Japan