1. Department of Physiology, Anatomy and Genetics, University of Oxford,South Parks Road, Oxford, UK
2. Department of Comparative Biomedical Sciences, Royal Veterinary College,Royal College Street, London NW1 0TU, UK
3. Clinical Biomanufacturing Facility, Nuffield Department of Clinical Medicine, University of Oxford, Old Road, Oxford, UK
4. Medical Research Council, Laboratory of Molecular Biology, Francis Crick Avenue, Cambridge, UK