1. Department of Biochemistry, Genetics and Microbiology, Forestry and Agricultural Biotechnology Institute (FABI), University of Pretoria, Private Bag X20, Pretoria 0028, South Africa
2. Mondi South Africa, Forests Operations, Research and Development Department, Trahar Technology Centre—TTC, PO Box 12, Hilton 3245, South Africa
3. Department of Forestry and Environmental Resources, College of Natural Resources, North Carolina State University, 2820 Faucette Drive, Raleigh, NC, USA