1. Department of Biological Sciences, Graduate School of Science, The University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo, 113-0033 Japan
2. Department of Biological Sciences, Osaka University, 1-1 Machikaneyama-cho, Toyonaka, Osaka, 560-0043 Japan
3. Department of Applied Biological Science, Faculty of Science and Technology, Tokyo University of Science, 2641 Yamazaki, Noda, Chiba, 278-8510 Japan
4. Division of Cellular Dynamics, National Institute for Basic Biology, 38 Nishigounaka, Myodaiji, Okazaki, Aichi, 444-8585 Japan
5. Department of Basic Biology, The Graduate University for Advanced Studies (SOKENDAI), 38 Nishigounaka, Myodaiji, Okazaki, Aichi, 444-8585 Japan