1. Department of Livestock and One Health, Institute of Infection, Veterinary and Ecological Sciences, University of Liverpool , Leahurst Campus, Wirral , UK
2. School of Veterinary Medicine and Science, University of Nottingham , Sutton Bonington Campus, Leicestershire , UK
3. Royal (Dick) School of Veterinary Studies and The Roslin Institute, University of Edinburgh , Easter Bush Campus, Midlothian , UK