1. Institute of Microbiology and Infection, School of Biosciences, University of Birmingham , Birmingham, B15 2TT , UK
2. Kent Fungal Group, Division of Natural Sciences, School of Biosciences, University of Kent , Canterbury, CT2 7NJ , UK
3. Institute of Microbiology and Infection, College of Medical and Dental Sciences, University of Birmingham , Birmingham, B15 2TT , UK