1. Department of Biological Sciences, University of Pittsburgh, Pittsburgh, PA, USA
2. Estación Biológica de Doñana, Consejo Superior de Investigaciones Científicas (EBD-CSIC), Sevilla, Spain
3. Department of Tropical Ecology, University of Yucatan, Mérida, Yucatán, México
4. Department of Biological Sciences, East Tennessee State University, Johnson City, TN, USA