1. Department of Genetics, Yale University School of Medicine, New Haven, CT 06510, USA
2. Department of Cell and Developmental Biology, Vanderbilt University School of Medicine, Nashville, TN 37232, USA
3. Department of Neuroscience, Yale University School of Medicine, New Haven, CT 06510, USA
4. Program in Neuroscience, Vanderbilt University School of Medicine, Nashville, TN 37232, USA