1. Plant Pathology and Plant-Microbe Biology Section, School of Integrative Plant Science, Cornell University , Geneva, NY 14456 , USA
2. Boyce Thompson Institute, Cornell University , Ithaca, NY 14853 , USA
3. Plant Biology Section, School of Integrative Plant Science, Cornell University , Ithaca, NY 14853 , USA
4. USDA-ARS Robert W. Holley Center for Agriculture and Health , Ithaca, NY 14853 , USA