1. Department of Ecology and Evolutionary Biology, Brown University, Providence, RI 02912, USA
2. Institute at Brown for Environment and Society, Brown University, Providence, RI 02912, USA
3. Department of Ecology and Evolutionary Biology, University of California—Los Angeles, Los Angeles, CA 90095, USA
4. Department of Biological Sciences and Museum of Natural Science, Louisiana State University, Baton Rouge, LA 70803, USA