1. Centre for Novel Agricultural Products, Department of Biology, University of York , York YO10 5DD , UK
2. Bioscience Technology Facility, Department of Biology, University of York , York YO10 5DD , UK
3. York Biomedical Research Institute, Department of Biology, University of York , York YO10 5DD , UK
4. Florida Museum of Natural History, University of Florida , Gainesville, FL 32611 , USA
5. School of Life Science, Yunnan Normal University , Kunming 650092, Yunnan , China
6. Key Laboratory of Yunnan for Biomass Energy and Biotechnology of Environment, Yunnan Normal University , Kunming 650500 , China
7. Key Laboratory for Plant Diversity and Biogeography of East Asia/Yunnan Key Laboratory for Integrative Conservation of Plant Species with Extremely Small Populations, Kunming Institute of Botany, Chinese Academy of Sciences , Kunming 650201 , China