1. Department of Biology, San Diego State University , San Diego, CA 92182 , USA
2. Department of Biology, Indiana University , Bloomington, IN 47408 , USA
3. National Institutes of Health , Bethesda, MD 20892 , USA
4. Department of Biological Sciences, California State University San Marcos , San Marcos, CA 92096 , USA
5. Department of Entomology, University of Kentucky , Lexington, KY 40506 , USA