1. National Engineering Laboratory for Rice and By-product Deep Processing, Central South University of Forestry and Technology , Changsha 410004 , P. R. China
2. Hunan Province, Key Laboratory of Plant Functional Genomics and Developmental Regulation, College of Biology, Hunan University , Changsha 410082 , P. R. China
3. State Key Laboratory of Hybrid Rice, Hunan Hybrid Rice Research Center, Hunan Academy of Agricultural Sciences , Changsha, 410125 , P. R. China