1. School of Biosciences, University of Nottingham , Sutton Bonington Campus, Loughborough , UK
2. Department of Plant Sciences, University of Cambridge , Cambridge , UK
3. School of Life Sciences, University of Essex , Colchester , UK
4. Institut de Recherche pour le Developpement , 911 Av. Agropolis, 34394 Montpellier , France
5. School of Biological Sciences, University of Aberdeen , Aberdeen , UK